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The study found that FOXO4 localizes predominantly in the cytoplasm of Leydig cells in young mice but translocates to the nucleus in aged animals the same FOXO4-p53 sequestration pattern seen in other senescent cell types

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10.1038/sj.gt.3303094 Gene Ther

These NAD+ deficits contribute to metabolic dysfunction by impairing insulin sensitivity, reducing mitochondrial function, and promoting inflammatory responses that exacerbate metabolic pathology in research models

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